Selected Publications


Ball, A. D., van Dijk, R. E., Lloyd, P., Pogány, Á, Dawson, D. A., Dorus, S. Bowie, R. C. K., Burke, T. & Szekely, T. (2017). Levels of extra-pair paternity are associated with parental care in penduline tits (Remizidae). Ibis 159: 449–455.

Bosque, C., Bosque, C. D. & Lloyd, P. (2017). Diet and time-activity budget of White-backed Mousebirds Colius colius in south-western South Africa. Ostrich 1–6.

Ewart, A., Popple, L. W. & Hill K. B. (2017). Five new species of grass cicadas in the genus Graminitigrina (Hemiptera: Cicadidae: Cicadettinae: Cicadettini) from Queensland and Northern Territory, Australia: comparative morphology, songs, behaviour and distributions. Zootaxa, 4228, 1–62.

Lloyd, P., Frauenknecht, B. D., du Plessis, M. A. & Martin, T. E. (2017). Comparative life history of the south temperate Cape Penduline Tit (Anthoscopus minutus) and north temperate Remizidae species. Journal of Ornithology 158: 569–577.  

Popple, L. W. (2017). A revision of the Ewartia oldfieldi (Distant) species complex (Hemiptera: Cicadidae: Cicadettinae) with five new species from eastern and northern Australia. Zootaxa 4263: 401–449.

Popple, L. W. & Emery, D. L. (2017 in press).  Two new species of Clinopsalta Moulds (Hemiptera: Cicadidae) and additional distribution records for Clinopsalta adelaida (Ashton), with notes on their distinctive calling songs. Records of the Australian Museum 69: 237–256.


Lloyd, P. & Martin, T. E. (2016). Fledgling survival increases with development time and adult survival across north and south temperate zones. Ibis 158: 135–143. 

Lloyd, P., Martin, T. E., Taylor, A., Braae, A. & Altwegg, R. (2016). Age, sex and social influences on adult survival in the co-operatively breeding Karoo scrub-robin. Emu 116: 394–401.

Popple, L. W. (2016). A new species of Macrotristria Stål from the Shoalwater Bay region of  central Queensland. Australian Entomologist 43: 101–107.

Popple, L. & Marshall, D. (2016). Australian cicadas: Worth a closer listen. Wildlife Australia 53: 24–26. 


Emery, N. J., Emery, D. L. & Popple, L. W. (2015). A redescription of Yoyetta landsboroughi (Distant) and Y. tristrigata (Goding and Froggatt) (Hemiptera: Cicadidae) and description of four new related species. Zootaxa 3948: 301–341.

Ewart, A., Popple, L. W. & Marshall, D. C. (2015). New species of Simona Moulds, 2012 and Chelapsalta Moulds, 2012 cicadas (Cicadidae: Cicadettinae: Cicadettini) from Australia: comparative morphology, songs, behaviour and distributions. Zootaxa 4001:  1-65.

Martin,T. E., Oteyza, J., Mitchell, A., Potticary, A. L. & Lloyd, P. (2015). Postnatal growth rates covary weakly with embryonic development rates and do not explain adult mortality probability among songbirds on four continents. American Naturalist 185: 380-389.

Popple, L. W. (2015). A new species of Toxala (Moulds) (Cicadidae: Cicadettinae: Cicadettini) from north Queensland, with notes on its calling song and comparisons with a congeneric species. Australian Entomologist 42: 77-90.

Searle, J. B., Prince, J. B., Stewart, D. & Lloyd, P. (2015). Breeding success of a subtropical Little Tern, Sternula albifrons sinensis, colony. Emu 116: 81–85.


Lloyd, P., Abadi, F., Altwegg, R. & Martin, T. E. (2014). South temperate birds have higher apparent adult survival than tropical birds in Africa. Journal of Avian Biology 45: 493–500. doi: 10.1111/jav.00454.

Ribeiro, Â. M., Lloyd, P., Dean, W. R. J., Brown, M. & Bowie, R. C. K. (2014). The Ecological and Geographic Context of Morphological and Genetic Divergence in an Understorey-Dwelling Bird. PLoS ONE 9: e85903. doi:10.1371/journal.pone.0085903.


Ewart, A. & Popple, L. W. (2013). Three new species of the genus Gudanga Distant (Cicadidae: Cicadettinae: Cicadettini) from Queensland; comparative morphology, songs, behaviour and distributions. Memoirs of the Queensland Museum 56: 355–406.

Ewart, A. & Popple, L. W. (2013). New species of Drymopsalta Heath Cicadas (Cicadidae: Cicadettinae: Cicadettini) from Queensland and Northern Territory, Australia, with overview of genus. Zootaxa 3520: 1–42.

Lloyd, P., Sanders, M., Reis, T. & Abbott, A. (2013). Targeted trapping surveys shed new light on the distribution and habitat characteristics of the Carpentarian pseudantechinus (Pseudantechinus mimulus), a threatened dasyurid marsupial. Australian Mammalogy

Noske, R.A., Mulyani, E. A. & Lloyd, P. (2013). Nesting beside old nests, but not over water, increases current nest survival in a tropical mangrove-dwelling warbler. Journal of Ornithology 154: 517-523.

Popple, L. W. (2013). A revision of the Pauropsalta annulata Goding & Froggatt species group (Hemiptera: Cicadidae) based on morphology, calling songs and ecology, with investigations into calling song structure, molecular phylogenetic relationships and a case of hybridisation between two subspecies. Zootaxa 3730: 1–102.


Ribeiro, A., Lloyd, P., Feldheim, K, Bowie, R. C. K. (2012). Microgeographic socio-genetic structure of an African cooperative breeding passerine revealed: integrating behavioural and genetic data. Molecular Ecology 21: 662–672.

Popple, L.W. (2012). An analysis of the calling song of Burbunga mouldsi Olive (Hemiptera: Cicadidae). Australian Entomologist 39: 75-78.

Petchey,  F., Ulm, S., David, B., McNiven, I.J., Asmussen, B., Tomkins, H., Richards, T., Rowe, C., Leavesley, M., Mandui, M. & Stanisic, J. 2012. 14C Marine reservoir variability in herbivores and deposit-feeding gastropods from an open coastline, Papua New Guinea. Radiocarbon 54: 1-12.

Shea, M., Colgan, D. & Stanisic, J. 2012. Systematics of the land snail genus Gyrocochlea and its relatives. (Mollusca: Charopidae). Zootaxa. 3585: 1-109.


Hugall, A. F. & Stanisic, J. 2011. Beyond the prolegomenon: a molecular phylogeny of the Australian camaenid land snail radiation. Zoological Journal of the Linnean Society 161: 531-572.

Lloyd, P., Weeks, S., Magno-Canto, M., Plagányi, G. & Plagányi, E.E. (2011). Ocean warming alters species abundance patterns and increases species diversity in an African sub-tropical reef-fish community. Fisheries Oceanography 21: 78–94.

Ribeiro, A., Lloyd, P., Feldheim, K, Bowie, R.C.K. (2011). A tight balance between natural selection and gene flow in a southern African arid-zone endemic bird. Evolution 65: 3499-3514.

Martin, T. E., Lloyd, P., Bosque, C., Barton, D. C., Biancucci, L., Cheng, Y.-R. and Ton, R. (2011). Growth rate variation among passerine species in tropical and temperate sites: an antagonistic interaction between parental food provisioning and nest predation risk. Evolution 65: 1607-1622.


Stanisic, J., Shea, M., Potter, D. and Griffiths, O. 2010. Australian Land Snails – Volume 1: A field guide to eastern Australian species. Bioculture Press, Mauritius.

Popple, L. W. and Emery, D. L. (2010). A new cicada genus and a redescription of Pauropsalta subolivacea Ashton (Hemiptera: Cicadidae) from eastern Australia. Australian Entomologist 37: 147-156.

Popple, L. W. and Walter, G. H. (2010). A spatial analysis of the ecology and morphology of cicadas in the Pauropsalta annulata species complex (Hemiptera: Cicadidae). Biological Journal of the Linnean Society 101: 553-565.

David, B., Geneste, J-M., Aplin, K., Delannoy, J-J., Araho, N., Clarkson, C., Connell, K., Haberle, S., Barker, B., Lamb, L., Stanisic, J. Fairbairn, A. & Skelly, R. 2010. The Emo Site (OAC), Gulf Province, Papua New Guinea:  resolving long-standing questions of antiquity and implications for the history of the ancestral hiri trade. Australian Archaeology 70: 1-30.


Lloyd, P., Taylor, W. A., du Plessis, M. A. and Martin, T. E. (2009). Females increase reproductive investment in response to helper-mediated improvements in allo-feeding, nest survival, nestling provisioning and post-fledging survival in the Karoo scrub-robin Cercotrichas coryphaeus.Journal of Avian Biology 40: 400-411.

Stanisic, J. 2009. Crikey steveirwini gen. et sp. nov. from montane habitats in the Wet Tropics of northeastern Queensland, Australia (Eupulmonata: Camaenidae). Zootaxa 2206: 62-68.


McNiven, I.J., Crouch, J., Weisler, M., Kemp, N., Martinez, L.C., Stanisic, J., Orr, M., Brady, L., Hocknull, S. & Boles, W. 2008. Tigershark Rockshelter (Baidamau Mudh): seascape and settlement reconfigurations on the Sacred Islet of Pulu, western Zenadh Kes (Torres Strait). Australian Archaeology 66: 15-32.

Stanisic, J. 2008. Land snails of Mua island, Torres Strait. Memoirs of the Queensland Museum, Cultural Heritage Series 4:1-4.

Osborn, A.W, Forteath, G.N.R. & Stanisic, J. 2008. A new species of freshwater sponge (Porifera:Spongillidae) of the genus Radiospongilla from lake Pedder in Tasmania. Papers and Proceedings of the Royal Society of Tasmania 142: 39-44.

Lloyd, P. (2008). Adult survival, dispersal and mate fidelity in the White-fronted Plover Charadrius marginatus. Ibis 150: 182-187.

Popple, L. W., Walter, G. H. and Raghu (2008). The structure of calling songs in the cicada Pauropsalta annulata Goding and Froggatt (Hemiptera: Cicadidae): evidence of diverging populations? Evolutionary Ecology 22: 203-215.


Baudains, T. P. and Lloyd, P. (2007). Habituation and habitat changes can moderate the impacts of human disturbance on shorebird breeding performance. Animal Conservation 10: 400-407.

David, B., Fairbairn, A., Aplin, K., Green, M., Stanisic, J., Weisler, M., Murepe, L., Simala, D., Kokents, T. & Dop, J. 2007. OJP, a terminal Pleistocene site in the Gulf Province lowlands, Papua New Guinea. Archaeology in Oceania 42: 31-33.

Delport, W., Crowe, T.M., Lloyd, P. & Bloomer, P. (2007). Population growth complicates the inference of maternal gene flow between populations of the nomadic and partially migratory Namaqua Sandgrouse (Pterocles namaqua). Journal of Heredity 98: 158-164.

Martin, T. E., Auer, S. K., Bassar, R.D., Niklison, A. and Lloyd, P. (2007). Geographic variation in avian incubation periods and parental influences on embryonic temperature. Evolution 61: 2558-2569.

Lloyd, P. (2007). Predator control, mesopredator release, and impacts on bird nesting success: a field test. African Zoology 42: 180-186.

Stanisic, J., Cameron, R.A.D., Pokryszko, B.M. & Nekola, J.C. 2007. Forest snail faunas from south-east Queensland and north-east New South Wales (Australia): patterns of local and regional richness and differentiation. Malacologia 49: 445-462.


Chambers, J., Williamson, I. and Wilson, J. C. (2006). Soil pH influences embryonic survival in Pseudophryne bibronii. Austral Ecology 31: 68-75.

Martin, T.E., Bassar, R.D., Auer, S.K., Fontaine, J.J., Lloyd, P., Mathewson, H. Chalfoun, A. & Niklison, A. (2006). Life-history and ecological correlates of geographic variation in egg and clutch mass among passerine species. Evolution 60: 390-398.

Lloyd, P. (2006). Density-dependent nest predation: a field test. African Journal of Ecology 44: 293-295.

Lloyd, P. (2006). Book review: Monitoring bird populations using mistnets. Ostrich: 77: 114-115.

Lloyd, P., Martin, T. E., Redmond, R. L., Hart, M. M., Langner, U. & Bassar, R. D. (2006). Assessing the influence of spatial scale on the relationship between avian nesting success and forest fragmentation: a case study. In Wu, J. B., Jones, B., Li, H. and Loucks, O. L. (eds). Scaling and Uncertainty Analysis in Ecology. Columbia University Press. pp. 255-269.


Emery, D. L., Emery, S. J., Emery, N. J. and Popple, L. W. (2005). A phenological study of the cicadas (Hemiptera: Cicadidae) in western Sydney, New South Wales, with notes on plant associations. Australian Entomologist 32: 97-110.

Hyman, I.T. & Stanisic, J. 2005. New charopid land snails chiefly from limestone outcrops in NSW (Eupulmonata: Charopidae). Memoirs of the Queensland Museum 50: 219-302.

Lloyd, P., Martin, T.E., Redmond, R.L., Langner, U. & Hart, M.M. (2005). Habitat fragmentation affects ecological processes and bird population viability. Ecological Applications 15: 1504-1514.

Stanisic, J., Burwell, C., Raven, R., Monteith, G. and Baehr, B. (2005). Terrestrial Invertebrate Status Review – Brisbane City. Brisbane City Council, Brisbane.


Stanisic, J. and Ponder, W. F. (2004). “Forest snails in eastern Australia.” In Lunney, D. (ed.). Conservation of Australia's forest fauna. Royal Zoological Society: Mosman. pp. 127-149.


Hugall, A., Stanisic, J. and Moritz, C. (2003). “Phylogeography of terrestrial gastropods: the case of the Sphaerospira lineage and history of Queensland rainforests.” In Lydeard, C. and Lindberg, D. (eds). Molecular systematics and phylogeography of molluscs. Smithsonian Books, Washington. pp. 270-301.

Popple, L. W. (2003). A new species of Cicadetta Amyot (Hemiptera: Cicadidae) from Queensland, with notes on its calling song. Australian Entomologist 30: 107-114.


Hugall, A., Moritz, C., Moussalli, A. and Stanisic, J. (2002). Reconciling palaeodistribution models and comparative phylogeography in the Wet Tropics rainforest land snail Gnarosophia bellendenkerensis (Brazier, 1875). Proceedings of the National Academy of Sciences of the United States of America 99: 6112-6117.

Popple, L. (2002). “Butterflies (Lepidoptera).” In Horton, H. (ed.). A Brisbane Bushland: the history and natural history of Enoggera Reservoir and its environs. Queensland Naturalists Club. pp. 101-106.

Popple, L. and Ewart, A. (2002). “Cicadas (Hemiptera).” In Horton, H. (ed.). A Brisbane Bushland: the history and natural history of Enoggera Reservoir and its environs. Queensland Naturalists Club. pp. 107-116.

Popple, L. W. and Strange A. D. (2002). Cicadas, and their songs, from the Tara and Warroo Shires, southern central Queensland.Queensland Naturalist 40: 15-30.


Popple, L. W. (2001). Bird observations in the Charleville, Quilpie and Windorah districts, southwestern Queensland. Queensland Naturalist 39: 72-79.

Moritz, C., Richardson, K.S., Ferrier, S., Monteith, G., Stanisic, J., Williams, S.E. & Whiffin, T. 2001. Biogeographical concordance and efficiency of taxon indicators for establishing conservation priority in a tropical rainforest biota. Proceedings of the Royal Society of London B. 268: 1875-1881.

Rowe, C., Stanisic, J., David, B. & Lourandos, H. 2001. The helicinid land snail Pleuropoma extincta (Odhner, 1917) as an environmental indicator in archaeology. Memoirs of the Queensland Museum 46: 741-770.


Stanisic, J. (2000). “Land snails.” In Ryan, M. (ed.), Wildlife of Tropical North Queensland. Queensland Museum, Brisbane. pp. 9-19.